1.   But biologists were not sure that the wolves had left until they saw the Crystal Bench wolves on Saturday afternoon a mile and a half from the pen.

2.   Even the coyotes who live at the margins of wolf country have prospered, thanks to the leftovers the wolves leave behind.

3.   If the wolves leave the pen, they will interrupt a beam of light that triggers an alarm on National Park Service radios.

4.   Once the first wolves have left the area, the other eight wolves, held in two pens, will be released.

5.   The wolves left other marks.

6.   Those coyotes that survive, he said, usually on the edge of wolf habitat, are flourishing because of the carcasses the wolves leave behind.